‘Queen of the Zoo’: Gramma, tortoise who lived through two world wars, dies aged 141

Gramma came to San Diego Zoo from Bronx Zoo in 1928 or 1931 and lived through 20 US presidents

After more than a century of munching on her favorite foods of romaine lettuce and cactus fruit, beloved Galapagos tortoise Gramma, the oldest resident of the San Diego Zoo, has died.

Gramma was born in her native habitat and was estimated to be about 141 years old, zoo officials said. She died on 20 November.